| The
horned toads are a group of specialized lizards from
desert and dry areas of northern and central America.
The genus is highly specialized on ants as diet.

Phrynosoma
Wiegmann, 1828
Anota
Hallowell, 1852
(Type = Anota mcallii)
Batrachosoma Fitzinger, 1843
Doliosaurus Girard, 1858
Tapaja Oken, 1816
Tapaya Fitzinger, 1826
Tapayia Gray, 1825
Tropidogaster Fitzinger, 1843 |
|
Phrynosoma
asio Cope, 1864
Batrachosoma
asio (Cope, 1864)
Phrynosoma spinimentum Peters,
1874 |
Distribution:
Guatemala
and Mexico. |
Phrynosoma
braconnieri Duméril & Bocourt,
1870
Distribution:
Mexico.
|
Phrynosoma
cornutum (Harlan, 1825)
Agama
cornutum Harlan, 1825
Lacerta cornuta (Harlan, 1825)
Phrynosoma bufonium Wiegmann, 1828
Phrynosoma cornutum planiceps Hallowell,
1852
Phrynosoma harlanii Wiegmann, 1834
Phrynosoma planiceps Hallowell,
1852
Tapaya cornuta (Harlan, 1825)
Tropidogaster cornutus (Harlan,
1825) |
Distribution:
Mexico and
USA. |
Phrynosoma
coronatum (Blainville, 1835) coronatum
(Blainville, 1835)
Agama
coronata
Blainville, 1835 |
blainvillei
Gray, 1839
Phrynosoma
blainvillei Gray, 1839
Phrynosoma ochoterenai Cuesta
Terron, 1932 |
cerroense
(Stejneger, 1893)
Phrynosoma
cerroense (Stejneger, 1893) |
frontale
Van Denburgh, 1894
Phrynosoma
frontalis
Van Denburgh, 1894 |
jamesi
Schmidt, 1922
Phrynosoma
jamesi
Schmidt, 1922 |
schmidti
Barbour, 1921
Phrynosoma
nelsoni Schmidt, 1922
Phrynosoma schmidti Barbour, 1921 |
Distribution: Mexico
(blainvillei, cerroense, coronatum, jamesi,
schmidti ) and USA
(blainvillei, frontale). |
Phrynosoma
ditmarsi Stejneger, 1906
Distribution:
Mexico.
|
Phrynosoma
douglassi (Bell, 1833)
Agama
douglasi Bell, 1833
Phrynosoma douglasi douglasi (Bell,
1833)
Phrynosoma douglasi exilis Cope,
1871
Phrynosoma douglasii pygmaea Yarrow,
1883 |
Distribution:
Canada and
USA |
Phrynosoma
hernandesi Girard, 1858
Phrynosoma
brevirostre (Girard, 1858)
Phrynosoma douglassi brachycercum
Smith, 1942
Phrynosoma douglassi brevirostre
(Girard, 1858)
Phrynosoma douglassi hernandezi
Girard, 1858
Phrynosoma douglassi ornatissimum
(Girard, 1858)
Phrynosoma douglassi ornatum
Girard, 1858
Phrynosoma hernandezi Girard,
1858
Phrynosoma ornatissimum (Girard,
1858)
Phrynosoma ornatum Girard, 1858
Tapaya brevirostris Girard, 1858
Tapaya ornatissima Girard, 1858
|
Distribution:
Mexico
and USA.
|
Phrynosoma
mcalli (Hallowell, 1852)
Anota
mcalli Hallowell, 1852 |
Distribution:
Mexico
and USA.
|
Phrynosoma
modestum Girard in Baird &
Girard, 1852
Anota
modesta (Girard in Baird & Girard,
1852)
Doliosaurus modestus (Girard in
Baird & Girard, 1852) |
Distribution:
Mexico
and USA.
|
Phrynosoma
orbiculare (Linnaeus, 1758) orbiculare
(Linnaeus, 1758)
Lacerta
orbicularis Linnaeus, 1758
Phrynosoma orbiculare alticola
Davis, 1953
Phrynosoma orbicularis (Linnaeus,
1758)
Phrynosoma wiegmanni Gray, 1839
Tapaya orbicularis (Linnaeus,
1758)
Tapaya orbicularis longicaudatus Dugés,
1888 |
boucardi
(Duméril & Bocourt, 1870)
Phrynosoma
boucardi (Duméril & Bocourt,
1870)
Tapaya boucardi Duméril
& Bocourt, 1870 |
bradti
Horowitz, 1955
Phrynosoma
orbiculare durangoensis Horowitz,
1955 |
cortezi
(Duméril & Bocourt, 1870)
Tapaya
orbicularis var. cortezi
Duméril & Bocourt, 1870 |
dugesi
(Duméril & Bocourt, 1870)
Tapaya
dugesii Duméril & Bocourt,
1870
Tapaya orbicularis var. dugesi
Duméril & Bocourt, 1870 |
orientale
Horowitz, 1955
Distribution: Mexico.
|
Phrynosoma
platyrhinos Girard in Baird &
Girard, 1852 platyrhinos
Girard in Baird & Girard,
1852
calidiarum
(Cope, 1896)
Anota
calidiarum Cope, 1896 |
goodei
Stejneger, 1893
Phrynosoma
goodei Stejneger, 1893 |
Distribution: Mexico
(calidiarum, goodei) and USA
(calidiarum, platyrhinos). |
Phrynosoma
solare Gray, 1845
Phrynosoma
regale Girard, 1858
Phrynosoma solaris Gray, 1845 |
Distribution:
Mexico
and USA.
|
Phrynosoma
taurus Dugès, 1869
Distribution:
Mexico.
|
|
|
| Literature |
Alberts, A. 1995. Conservation of the San Diego coast
horned lizard on the Southwestern Riverside County Multi-Species
Reserve: foraging ecology, microhabitat use, genetics,
and reproductive biology. Agreement No. 1550 - Progress
Report, 1(12/95): 1-8. |
Alberts, A. 1997. Conservation of the San Diego coast
horned lizard on the Southwestern Riverside County Multi-Species
Reserve: foraging ecology, microhabitat use, genetics,
and reproductive biology. Agreement No. 1550 - Progress
Report, 2(3/97): 1-12. |
| Ashton,
K.G. & K.L. Ashton. 1998. Phrynosoma douglasi.
Reproduction. Herpetol. Rev., 29(3): 168-169. |
| Axtell,
R.W. 1986. The Epipterygoids in Phrynosoma solare,
Still Present, but Disappearing. J. Herpetol., 20(1):
79-81. |
Axtell, R.W. 1988. Phrynosoma modestum - Interpretive
atlas of Texas lizards, 6: 1-18. |
Axtell, R.W. 1996. Phrynosoma cornutum - Interpretive
atlas of Texas lizards, 16: 1-9. |
Baharav, D. 1971. Movement ecology of the horned lizard,
Phrynosoma solare. Unpubl. M.S. Thesis, Arizona
State Univ., 65 pp. |
Baharav, D. 1975. Movement of the horned lizard Phrynosoma
solare. Copeia, 1975(4): 649-657. |
| Baird,
S.F. & C. Girard. 1852. Characteristics of some New
Reptiles in the Museum of the Smithsonian Institution.
Proc. Acad. Nat. Sci. Philadelphia, 6: 68-70. |
Ballinger, R.E. 1974. Reproduction of the Texas horned
lizard, Phrynosoma cornutum. Herpetologica, 30(4):
321-327. |
Banta, B.H. 1961. Herbivorous feeding of Phrynosoma
platyrhinos in southern Nevada. Herpetologica, 17(2):
136-137. |
Banta, B.H. 1968. The systematic status of the horned
lizard inhabiting Cedros Island, off the west coast of
Baja California, Mexico - Abstract. Joint ann. meet. Amer.
asoc. adv. sci. & HL, Dallas, Texas. |
Barbour, T. 1921. A new Phrynosoma from Cerros
Island. proc. New England Zool. Club, 7: 113-115. |
Baur, B. 1973. Pflege und Zucht der Wüstenkrötenechse
Phrynosoma platyrhinos. Salamandra, 9(3/4): 145-159. |
Baur, B.E. 1974. Colima-Krötenechse (Phrynosoma
asio Cope 1864) - der mexikanische Riese. Das Aquarium,
58: 164-168. |
Baur, B. 1979. Leben in der Wüste: Krötenechsen
(Phrynosoma Wiegmann, 1828). 2. Teil: Pflege
und Zucht der Rundschwanz-Krötenechse Phrynosoma
modestum Girard, 1852. Das Aquarium, 125: 528-532.
|
Baur, B. 1979. Pflege und "Zucht" der Riesenkrötenechse
Phrynosoma asio. Salamandra, 15(1): 1-12. |
| Baur,
B. 1984. Krötenechsen-Bastarde (Phrynosoma).
Salamandra, 20(2/3): 70-87. |
Baur, B.E. 1986. Die Felsen-Krötenechse (Phrynosoma
ditmarsi Stejneger, 1906). herpetofauna, 8(40): 27. |
Baur, B.E. 1986. Longevity of horned lizards of the genus
Phrynosoma. Bull. Maryland Herpetol. Soc., 22(3):
149-151. |
Baur, B.E. 1987. Kurzhorn-Krötenechsen. Phrynosoma
douglassi and Phrynosoma orbiuculare. Teil
1. Das Aquarium, 217: 421-426. |
Baur, B.E. 1987. Kurzhorn-Krötenechsen. Phrynosoma
douglassi and Phrynosoma orbiuculare. Teil
2. Das Aquarium, 219: 476-478. |
Baur, B.E. 1988. Die Wüsten-Krötenechse Phrynosoma
platyrhinos. Teil 1: Verbreitung - Aussehen - Ernährung.
Das Aquarium, 225: 154-158. |
Baur, B.E. 1988. Die Wüsten-Krötenechse Phrynosoma
platyrhinos. Teil 2: Überwinterung - Fortpflanzung
- Systematik. Das Aquarium, 227: 303-306. |
| Baur,
B. & R.R. Montanucci. 1998. Krötenechsen. Lebensweise.
Pflege. Zucht. Herpeton, Offenbach, 158 pp. |
| Behler,
J.L. & F.W. King. 1979. The Audubon Society Field
Guide to North American Reptiles & Amphibians. Alfred
A. Knopf, New York, 743 pp. |
Bell, T. 1828. The description of a new species of Agama,
brought from the Columbia River by Mr. Douglas. Trans.
Linn. Soc. London, 16(1): 105-107. |
| Bergman,
E., B. Hill, C. Montgomery, T. Childers, J.D. Manzer,
J. Sifert & S.P. Mackessy. 1998. Geographic Distribution.
Phrynosoma douglasii. Herpetol. Rev., 29(2):
111. |
| Beauchamp,
B., B. Wone, S. Bros & M. Kutilek. 1998. Habitat Use
of the Flat-tailed Horned Lizard (Phrynosoma mcallii)
in a Disturbed Environment. J. Herpetol., 32(2): 210-216. |
Blaney, R.M. & P.J. Kimmich. 1973. Notes on the Young
of the Texas Horned Lizard, Phrynosoma cornutum.
HISS News J., 1(4): 120. |
Blount, R.F. 1929. Seasonal cycles of the intestitial
cells in the testes of the horned toad (Phrynosoma
solare). Seasonal variations in the number and morphology
of the interstitial cells and the volume of the interstitial
tissue. J. Morphol., 48(2): 317-344. |
| Bott,
K., T. Brown & D. Eifler. 2001. Phrynosoma cornutum.
Diet. Herpetol. Rev., 32(1): 41. |
Boulenger, E.G. 1916. On a new lizard of the genus Phrynosoma,
recently living in the society's gardens. Proc. Zool.
Soc. London, 1916: 537-538. |
| Brattstrom,
B.H. 1997. Status of the Subspecies of the Coast Horned
Lizard, Phrynosoma coronatum. J. Herpetol., 31(3):
434-436. |
Brown, T.L. & R.V. Lucchino. 1972. A record sized
specimen of the Texas horned lizard (Phrynosoma cornutum).
Texas J. Sci., 14(3): 353-354. |
Bryant, H.C. 1911. The horned lizards of California and
Nevada of the genera Phrynosoma and Anota.
Univ. California Publ. Zool., 9(1): 1-84. |
Bundy, R.E. & J. Neess. 1958. Color variation in the
round-tailed horned lizard Phrynosoma modestum.
Ecology, 39(3): 463-477. |
Burleson, G.L. 1942. The source of the blood ejected from
the eye by horned toads. Copeia, 1942(4): 246-248. |
Burrow, A.L, R.T. Kazmair, E.C. Hellgren & D.C. Ruthven.
2001. Microhabitat selection by Texas Horned Lizards in
southern Texas. J. Wildlife Management, 65: 645-652. |
Cohen, A.c. & J.L. Cohen. 1990. Ingestion of blister
beetles by a Texas horned lizard. Southwestern Nat., 35(3):
369. |
Collins, A. 1988. Natural history of the desert horned
lizard Phrynosoma platyrhinos in the central
Mojave Desert. Proc. Conf. California Herpet., 1988: 29-37. |
Cope, E.D. 1871. Phrynosoma douglassii exilis.
Ann. Rept. U. S. Geol. Surv. Terrs., 1871: 46. |
Cutter, W.L. 1959. An instance of blood-squirting by Phrynosoma
solare. Copeia, 1959(2): 176. |
Dammann, J. 1949. Birth of 18 young Phrynosoma douglassii
hernandesi (Girard). herpetologica, 5: 144. |
Davis, W.B. 1953. A new horned lizard, genus Phrynosoma,
from Mexico. Proc. Biol. Soc. Washington, 66: 27-30. |
Donaldson, W., A.H. Price & J. Morse. 1994. The current
status and future prospects of the Texas horned lizard
(Phrynosoma cornutum) in Texas. Texas J. Sci.,
46(2): 97-113. |
Dugés, A. 1869. Catalogo de animales vertebrados
observados en la republica Mexicana. La Naturaleza, 1:
137-145. |
Dumas, P.C. 1964. Species-pair allopatry in the genera
Rana and Phrynosoma. Ecology, 45(1):
178-181. |
Duncan, R.B., T.C. Esque & K.L. Echols. 1994. Phrynosoma
mcallii. Predation. Herpetol. Rev., 25(2): 68. |
Edgren, R.A., Jr. 1948. Notes on a large specimen of Phrynosoma
solare Gray. Copeia, 1948(2): 129. |
| Fair,
W.S. & S.E. Henke. 1997. Efficacy of Capture Methods
for a Low Density Population of Phrynosoma cornutum.
Herpetol. Rev., 28(3): 135-137. |
| Fair,
W.S. & S.E. Henke. 1999. Movements, Home Ranges, and
Survival of Texas Horned Lizards (Phrynosoma cornutum).
J. Herpetol., 33(4): 517-525. |
| Fisher,
M. & A. Muth. 1995. A Backpack Method for Mounting
Radio Transmitters to Small Lizards. Herpetol. Rev., 26(3):
139-140. |
Funk, R.S. 1981. Phrynosoma mcalli. Cat. Amer.
Amphib. Rept., 281: 1-2. |
| Gentry,
A.F. 1885. A Review of the Genus Phrynosoma.
Proc. Acad. Nat. Sci. Philadelphia, (3)37: 138-148. |
Girard, C.F. 1852. A monographic essay on the genus Phrynosoma.
Stansb. Expl. surv. Great Salt Lake of Utah, 1852: 354-365. |
Girard, C. 1858. Herpetology. in United States
Exploring Expedition during the years 1838, 1839, 1840,
1841, 1842. Under the command of Charles Wilkes, U.S.N.
J.B. Lippincott and Co., Phladelphia. Vol. 20: xviii +
496 pp. |
Goldberg, S.R. 1971. Reproduction in the short-horned
lizard Phrynosoma douglassi in Arizona. Herpetologica,
27(3): 311-314. |
Goldberg, S.R. 1983. Reproduction of the coast horned
lizard Phrynosoma coronatum in southern California,
USA. Southwestern. Nat., 28(4): 478-479. |
| Goldberg,
S.R. 1999. Phrynosoma ditmarsi. Testis Cycle.
Herpetol. Rev., 30(3): 167. |
Goldberg, S.R. & C.R. Bursey. 1991. Gastrointestinal
helminths of the Mexican horned lizards Phrynosoma
braconnieri and Phrynosoma taurus (Iguanidae).
Southwestn. Nat., 36(3): 365-368. |
| Goldberg,
S.R. & C.R. Bursey. 2000. Phrynosoma ditmarsi.
Endoparasites. Herpetol. Rev., 31(4): 242. |
Goldberg, S.R., C.R. Bursey & R. Tawil. 1993. Gastrointestinal
helminths of five horned lizard species Phrynosoma
(Phrynosomatidae) from arizona. J. Helminth. Soc. Washington,
60(2): 234-238. |
| Grant,
T.D. & A.C. Alberts. Phrynosoma coronatum.
Predation and Telemetry. Herpetol. Rev., 32(4): 257. |
Gray, J.E. 1845. Catalogue of the specimens of lizards
in the collection of the British Museum. Edward Newman,
London, xxvi + 289 pp. |
| Grismer,
L.L. & E. Mellink. 1994. The Addition of Sceloporus
occidentalis to the Herpetofauna of Isla de Cedros,
Baja California, México and its Historical and
Taxonomic Implications. J. Herpetol., 28(1): 120-126. |
Guerra, T. 1997. Genetic analysis of the texas horned
lizard, Phrynosoma cornutum - Horned Lizard Conservation
Society. Third national horned lizard conference. |
Guyer, C. & A.D. Linder. 1985. Growth and population
structure of the short-horned lizard Phrynosoma douglassi
and the sagebrush lizard Sceloporus graciosus
in southeastern Idaho. Northw. Sci., 59(4): 294-303. |
Guyer, C. & A.D. Linder. 1985. Thermal ecology and
activity patterns of the short-horned lizard Phrynosoma
douglassi and the sagebrush lizard Sceloporus
graciosus in southeastern Idaho. Gt. Basin Nat.,
45(4): 607-614. |
| Hager,
S.B. 1995. Phrynosoma coronatum blainvillii.
Growth. Herpetol. Rev., 26(3): 145-146. |
| Hager,
S.B. 1996. Phrynosoma coronatum blainvillii.
Growth. Herpetol. Rev., 27(2): 80. |
| Hallowell,
E. 1852. Descriptions of new Species of Reptiles inhabiting
North America. Proc. Acad. Nat. Sci. Philadelphia, 6:
177-182. |
Hammerson, G.A. & H.M. Smith. 1991. The correct spelling
of the name for the short-horned lizard of North America.
Bull. Maryland Herpetol. Soc., 27(3): 121-127. |
Harlan, R. 1824/1825. Description of two new species of
Agama. J. Acad. Nat. Sci. Philadelphia, 4(2):
296-304. |
Hazen-Hammond, S. 1994. Here's to the "horny toad",
one tough little squirt. "Horny toads" enjoy
a special place in western hearts. Smithsonian, 25(9):
82-91. |
Heath, J.E. 1965. Temperature regulation and diurnal activity
in horned lizards. Univ. California Publ. Zool., 64(3):
97-136. |
| Henke,
S.E. 1998. The Effect of Multiple Search Items and Item
Abundance on the Efficiency of Human Searchers. J. Herpetol.,
32(1): 112-115. |
| Henke,
S.E. & M. Montemayor. 1997. Phrynosoma cornutum.
Growth. Herpetol. Rev., 28(3): 152. |
| Henke,
S.E. & M. Montemayor. 1998. Diel and Monthly Variations
in Capture Success of Phrynosoma cornutum via
Road Cruising in Southern Texas. Herpetol. Rev., 29(3):
148-150. |
| Hernández-Ibarra,
X., A. Ramirez-Bautista & R. Torres-Cervantes. 2000.
Phrynosoma orbiculare orbiculare. Reproduction.
Herpetol. Rev., 31(3): 175-176. |
Hewatt, W.G. 1937. Courting and egg-laying habits of Phrynosoma
cornutum. Copeia, 1937(4): 234. |
| Hodges,
W.L. 1995. Phrynosoma ditmarsi. Cat. Amer. Amphib.
Rept., 614: 1-3. |
Horowitz, S.B. 1955. An arrangment of the subspecies of
the horned toad, Phrynosoma orbiculare (Iguanidae).
Amer. Midl. Nat., 54(1): 204-218. |
Howard, C.W. 1974. Comparative reproductive ecology of
horned lizard (genus Phrynosoma) in southwestern
USA and northern Mexico. J. Arizona Acad. Sci., 9(3):
108-116. |
International Commission on Zoological Nomenclature. 1959.
Opinion 565: Validation under the plenary powers of the
generic name "Phrynosoma" Wiegmann,
1828 (Class Reptilia) ("Opinion" supplementary
to "Opinion" 92). Opin. Decl. Int. Commn. Zool.
Nom., 20(32): 359-376. |
Jennings, M.R. 1985. Correction of the type-locality of
Phrynosoma coronatum schmidti. Herpetol. Rev.,
16(4): 107. |
| Jennings,
M.R. 1987. Impact of the Curio Trade for San Diego Horned
Lizards (Phrynosoma coronatum blainvillii) in
the Los Angeles Basin, California: 1885-1930. J. Herpetol.,
21(4): 356-358. |
Jennings, M.R. Phrynosoma cerroense. Cat. Amer.
Amphib. Rept., 427: 1-2. |
Jennings, M.R. Phrynosoma coronatum. Cat. Amer.
Amphib. Rept., 428: 1-5. |
Klauber, L.M. 1932. The flat-tailed horned toad in Lower
California. Copeia, 1932(2): 100. |
Koenig, L. 1960. Zur Jugendentwicklung der Krötenechse
Phrynosoma douglassii hernandesi. Die Pyramide,
1: 15-17. |
| LaBonte,
J.P. 2001. Phrynosoma coronatum. Predation and
Telemetry. Herpetol. Rev., 32(4): 257-258. |
Laird, M. & R. Leech. 1980. Observations on the short-horned
lizard in southeastern Alberta. Blue Jay, 38(4): 214-218. |
Lawrence, J. & D. Wilhoft. 1958. Cryptic coloration
in lava-dwelling horned lizards. Copeia, 1958(1): 43-44. |
| Leatherman,
B.M. 1996. Phrynosoma coronatum blainvillii.
Conservation. Herpetol. Rev., 27(2): 80. |
| Lemos-Espinal,
J.A., D. Chiszar, C. Henke & H.M. Smith. 1998. Phrynosoma
cornutum. Predation. Herpetol. Rev., 29(3): 168. |
| Lemos-Espinal,
J.A., H.M. Smith & D. Chiszar. 2001. Phrynosoma.
Protection. Herpetol. Rev., 32(1): 41-42. |
Lowe, C.H., Jr. 1954. Normal field movements and growth
rates of marked regal horned lizards (Phrynosoma solare).
Ecology, 35(3): 420-421. |
Lowe, C.H. & C.W. Howard. 1975. Viviparity and reproductive
pattern in Phrynosoma ditmarsi in Sonora Mexico.
Southwestn. Nat., 20(2): 265-270. |
Lowe, C.H., M.D. Robinson & V.D. Roth. 1971. A population
of Phrynosoma ditmarsi from Sonora, Mexico. J.
Arizona Acad. Sci., 6(4): 275-277. |
Lynn, R.T. 1965. A comparative study of display behaviour
in Phrynosoma (Iguanidae). Southwestn. Nat.,
10(1): 25-30. |
Manaster, J. 1997. Horned lizards. Univ. Texas Press,
81 pp. |
Mayhew, W.W. 1965. Hibernation in the horned lizard Phrynosoma
m'calli. Comp. biochem. physiol., 16: 103-119. |
Mayhew, W.W. & S.J. Wright. 1971. Water impermeable
skin of the lizard Phrynosoma mcalli. Herpetologica,
27(1): 8-11. |
| Mayne,
P.J. 1997. Geographic Distribution. Phrynosoma modestum.
Herpetol. Rev., 28(1): 50. |
Medica, P.A., F.B. Turner & D.D. Smith. 1973. Effects
of radiation on a fenced population of horned lizards
Phrynosoma platyrhinos in southern Nevada. J.
Herpetol., 7(2): 79-85. |
Mendez, R.A. 1992. Seeing red. Why does the horned toad
cry tears of blood? Nat. Hist., 10: 48-49. |
Middendorf, G.A. III & W.C. Sherbrooke. 1992. Canid
elicitation of blood-squirting in a horned lizard (Phrynosoma
cornutum). Copeia, 1992(2): 519-527. |
Milne, L.J. 1938. Mating of Phrynosoma cornutum.
Copeia, 1938(4): 200. |
Milroy, L.G. III. 1997. Ebb and flow of populations of
the San Diego coast horned lizard - Horned Lizard Conservation
Society. Third national horned lizard conference. |
Minke, E.H. 1915. Die Krötenechse Phrynosoma
cornutum. Bl. Aquar.-u. Terrarienk., 26: 132-134.
|
Modry, D. 1993. Rozmnozeni ropusniku Phrynosoma orbiculare
cortezi v terariu. Akvarium a Terarium, 37(6): 38-43. |
Montanucci, R.R. 1979. Notes on systematics of horned
lizards allied to Phrynosoma orbiculare (Lacertilia;
Iguanidae). Herpetologica, 35(2): 116-124. |
Montanucci, R.R. 1981. Habitat separation between Phrynosoma
douglassi and Phrynosoma orbiculare (Lacertilia;
Iguanidae) in Mexico. Copeia, 1981(1): 147-153. |
Montanucci, R.R. 1983. Breeding, captive care and longevity
of the short-horned lizard Phrynosoma douglassi.
Int. Zoo Yb., 23: 148-156. |
Montanucci, R.R. 1987. A phylogenetic study of the horned
lizards, genus Phrynosoma, based on skeletal
and external morphology. Contr. Sci., Los Angeles Co.
Mus., 390: 1-36. |
Montanucci, R.R. 1989. Maintenance and propagation of
horned lizards (Phrynosoma) in captivity. Bull.
Chicago Herpetol. Soc., 24(12): 229-238. |
| Montanucci,
R.R. 1989. The Relationship of Morphology to Diet in the
Horned Lizard Genus Phrynosoma. Herpetologica,
45(2): 208-216. |
Montanucci, R.R. 1989. Unique behaviors in captive rock
horned lizards, Phrynsoma ditmarsi. Herpetol.
Rev., 20(4): 85-87. |
Montanucci, R.R. 1989. The reproduction and growth of
Phrynosoma ditmarsi (Sauria: Iguanidae) in captivity.
Zoo Biol., 8: 139-149. |
Montanucci, R.R. 1996. Morphological variation in the
gular fold in the horned lizard genus, Phrynosoma
(Iguania: Phrynosomatidae). Herpetologica, 52(1):
46-55. |
Montanucci, R.R. & B.E. Baur. 1982. Mating and courtship
related behaviors of the short-horned lizard Phrynosoma
douglassi. Copeia, 1982(4): 971-974. |
| Montgomery,
C., T. Childers, J.D. Manzer, E. Bergman, J. Sifert, B.
Hill & S.P. Mackessy. 1998. Geographic Distribution.
Phrynosoma cornutum. Herpetol. Rev., 29(2): 110. |
| Montgomery,
C. & S.P. Mackessy. 2001. Phrynosoma cornutum.
Movement. Herpetol. Rev., 32(4): 258. |
| Montgomery,
C.E., S.P. Mackessy & J.C. Moore. 2003. Body Size
Variation in the Texas Horned Lizard, Phrynosoma cornutum,
from Central Mexico to Colorado. J. Herpetol., 37(3):
550-553. |
Munger, J.C. 1984. Home ranges of horned lizards (Phrynosoma):
circumscribed and exclusive? Oecologia, 62: 351-360. |
Munger, J.C. 1984. Long-term yield from harvester ant
colonies: implications for horned lizard foraging strategy.
Ecology, 65(4): 1077-1086. |
Munger, J.C. 1984. Optimal foraging? Patch use by horned
lizards (Iguanidae: Phrynosoma). Am. Nat., 123(5):
654-680. |
Munger, J.C. 1986. Rate of death due to predation for
two species of horned lizard, Phrynosoma cornutum
and Phrynsoma modestum. Copeia, 1986(3):
820-824. |
Norris, K.S. 1949. Observations on the habits of the horned
lizard Phrynsoma m'callii. Copeia, 1949(3: 176-180. |
Norris, K.S. & C.H. Lowe, Jr. 1951. A study of the
osteology and musculature of Phrynosoma m'callii
pertinent to its systematic status. Bull. Chicago Acad.
Sci., 9(7): 117-125. |
Parker, W.S. 1971. Ecological observations on the regal
horned lizard (Phrynosoma solare) in Arizona.
Herpetologica, 27(3): 333-338. |
| Parker,
W.S. 1974. Phrynosoma solare. Cat. Amer. Amphib.
Rept., 162: 1-2. |
Perrill, R.H. 1983. Geogrpahic Distribution. Phrynosoma
ditmarsi. Herpetol. Rev., 14(4): 123. |
| Peters,
W.C.H. 1874. Über neue Saurier (Spæriodactylus,
Anolis, Phrynosoma, Tropidolepisma, Lygosoma, Ophioscincus)
aus Centralamerica, Mexico, und Australien. Mber. Königl.
Akad. Wiss. Berlin, 1873: 738-747. |
| Phillips,
J.A. & H.J. Harlow. 1981. Elevation of Upper Voluntary
Temperatures after Shielding the Parietal Eye of Horned
Lizards (Phrynosoma douglassi). Herpetologica,
37(4): 199-205. |
Pianka, E.R. 1991. Phrynosoma platyrhinos. Cat.
Amer. Amphib. Rept., 517: 1-4. |
| Pianka,
E.R. & W.S. Parker. 1975. Ecology of Horned Lizards:
A Review with Special Reference to Phrynosoma platyrhinos.
Copeia, 1975(1): 141-162. |
Porter, T. 1982. Phrynosoma coronatum jamesi. Longevity.
Herpetol. Rev., 13(1): 19. |
Powell, G.L. 1982. The eastern short-horned lizard in
Alberta: basic field ecology of northern marginal populations.
Diplomarbeit: Univ. Calgary, 343 pp. |
Powell, G.L. & A.P. Russell. 1984. The diet of the
eastern short-horned lizard (Phrynosoma douglassi
brevirostre) in Alberta and its relationship to sexual
size dimorphism. Can. J. Zool., 62(3): 428-440. |
Powell, G.L. & A.P. Russell. 1985. Field thermal ecology
of the eastern short-horned lizard (Phrynosoma douglassi
brevirostre). in southern Alberta. Can. J. Zool.,
63(2): 228-238. |
Powell, G.L. & A.P. Russell. 1985. Growth and sexual
size dimorphism in Alberta populations of the eastern
short-horned lizard, Phrynosoma douglassi brevirostre.
Can. J. zool., 63(1): 139-154. |
Powell, G.L. & A.P. Russell. 1991. Distribution of
the eastern short-horned lizard (Phrynosoma douglassi
brevirostre) in Alberta, Canada. Northw. Nat., 72(1):
21-26. |
Powell, G.L. & A.P. Russell. 1991. Parturition and
clutch characteristics of short-horned lizards (Phrynosoma
douglassi brevirostre) from Alberta. Can. J. Zool.,
69: 2759-2764. |
Powell, G.L. & A.P. Russell. 1998. The status of short-horned
lizards, Phrynosoma douglasli and P. hernandezi,
in Canada. Can. Fld. Nat., 112(1): 1-16. |
| Powell,
G.L., A.P. Russell & M.J. Ryan. 2002. Ontogenetic
Scaling of the Cranial Horn Array in Phrynosoma orbiculare
(Squamata: Phrynosomatidae). J. Herpetol., 36(4):
578-589. |
Presch, W. 1969. Evolutionary osteology and relationships
of the horned lizard genus Phrynosoma (Family
Iguanidae). Copeia, 1969(2): 250-275. |
Price, A.H. 1990. Phrynosoma cornutum. Cat. Amer.
Amphib. Rept., 469: 1-7. |
Prieto, A.A., Jr. & W.G. Whitford. 1971. Physiological
responses to temperature in the horned lizards, Phrynsoma
cornutum and Phrynosoma douglassi. Copeia,
1971(3): 498-504. |
Ramsey, L.W. 1956. Nesting of Texas horned lizards. Herpetologica,
12: 239-240. |
Reeder, T.W. 1995. Phylogenetic relationships among phrynosomatid
lizards as inferred from mitochondrial ribosomal DNA sequences:
substitution bias and information content of transitions
relative to transversions. Mol. Phylogenet. Evol., 4:
203-222. |
| Reeder,
T.W. &. R.R. Montanucci. 2001. Phylogenetic Analysis
of the Horned Lizards (Phrynosomatidae: Phrynosoma):
Evidence from Mitochondrial DNA and Morphology. Copeia,
2001(2): 309-323. |
Reeder, T.W. & J.J. Wiens. 1996. Evolution of the
lizard family Phrynosomatidae as inferred from diverse
types of data. Herpetol. Monogr., 10: 43-84. |
Reese, A.M. 1922. Egg-laying by the "horned toad",
Phrynosoma cornutum. Copeia, 1922: 15-16. |
Reeve, W.L. 1952. Taxonomy and distribution of the horned
lizards genus Phrynosoma. Univ. Kansas Sci. Bull.,
34(14): 817-960. |
| Richmond,
J.Q. 1998. Backpacks for Lizards: A Method For Attaching
Radio Transmitters. Herpetol. Rev., 29(4): 220-221. |
Rissing, S.W. 1981. prey preferences in the desert horned
lizard (Phrynsoma platyrhinos): influence of
prey foraging method and aggressive behavior. Ecology,
62(4): 1031-1040. |
| Rondeau,
R. 2002. Geographic Distribution. Phrynosoma modestum.
Herpetol. Rev., 33(3): 226. |
Rorabaugh, J.C., C.L. Palermo & S.C. Dunn. 1987. Distribution
and relative abundance of the flat-tailed horned lizard
(Phrynosoma mcallii) in Arizona. Southwestn.
Nat., 32(1): 103-109. |
Roth, V.D. 1971. Feed habits of ditmars horned lizard
with speculation on its type locality. J. Arizona Acad.
Sci., 6(4): 278-281. |
Roth, V.D. 1997. Ditmars' horned lizard (Phrynosoma
ditmarsi) or the case of the lost lizard. Sonoran
Herpetologist, 10(1): 2-6. |
Ruthling, P.D. 1919. Blood expelling of the horned lizards
in Mexico (Ph. orbiculare, Ph. taurus). Copeia,
72: 67-68. |
| Sánchez,
E.B., F.M. Quijano & W.C. Sherbrooke. 2000. Geographic
Distribution. Phrynosoma orbiculare. Herpetol.
Rev., 31(1): 54. |
| Sattler,
P.W. & J.S. Ries. 1995. Intraspecific Genetic Variation
among Four Populations of the Texas Horned Lizard, Phrynosoma
cornutum. J. Herpetol., 29(1): 137-141. |
Savage, J.M. 1954. Notulae herpetologicae 1-7. Trans.
Kansas Acad. Sci., 57(3): 326-334. |
Schmidt, P.J., W.C. Sherbrooke & J.O. Schmidt. 1989.
The detoxification of ant (Pogonomyrmex) venom
by a blood factor in horned lizards (Phrynosoma).
Copeia, 1989(3): 603-607. |
Schonberger, C.F. 1945. Oviposition by Phrynosoma
solare. Copeia, 1945(1): 53-54. |
Shaffer, D.t., Jr. & W.G. Whitford. 1981. Behavioral
resposnes of a predator, the round-tailed horned lizard,
Phrynosoma modestum and its prey, honey-pot ants,
Myrmecocystus spp. Am. Midl. Nat., 105(2): 209-216. |
Sheffield, S.R. & N. Carter. 1994. Phrynosoma
cornutum. Arboreal behavior. Herpetol. Rev., 25(2):
65-68. |
Sherbrooke, W.C. 1981. Horned lizards: unique reptiles
of western North America. Southwest Parks and Monuments
Assoc. globe, Arizona, 1-48. |
Sherbrooke, W.C. 1987. Captive Phrynosoma solare raised
without ants or hibernation. Herpetol. Rev., 18(1): 11-13. |
Sherbrooke, W.C. 1987. Defensive head posture in horned
lizards (Phrynosoma: Sauria: Iguanidae). Southwestn.
Nat., 32(4): 512-515. |
Sherbrooke, W.C. 1990. Predatory behavior of captive greater
roadrunners feeding on horned lizards. Wilson Bull., 102(1):
171-174. |
| Sherbrooke,
W.C. 1990. Rain-harvesting in the Lizard, Phrynosoma
cornutum: Behavior and Integumental Morphology. J.
Herpetol., 24(3): 302-308. |
Sherbrooke, W.C. 1991. Behavioral (predator-prey) interactions
of captive grasshopper mice (Onychomys torridus)
and horned lizards (Phrynosoma cornutum and P.
modestum). Amer. Midl. Nat., 126(1): 187-195.
|
| Sherbrooke,
W.C. 1995. Collecting and Feeding Harvester Ants (Pogonomyrmex
spp.) to Captive Horned Lizards (Phrynosoma spp.).
Herpetol. Rev., 26(1): 25-26. |
Sherbrooke, W.C. 1997. Ditmars' horned lizard, or rock
horned lizard: an historical update since rediscovery
(1970). Sonoran Herpetologist, 10(1): 6-8. |
Sherbrooke, W.C. 1997. Physiological (rapid) change of
color in horned lizards (Phrynosoma) of arid
habitats: hormonal regulation, effects of temperature,
and role in nature. Amphibia-Reptilia, 18: 155-175. |
| Sherbrooke,
W.C. 2000. Phrynosoma modestum. Nocturnal Nest-Digging
and Oviposition. Herpetol. Rev., 31(4): 242-243. |
| Sherbrooke,
W.C. 2002. Seasonally Skewed Sex-Ratios of Road-Collected
Texas Horned Lizards (Phrynsoma cornutum). Herpetol.
Rev., 33(1): 21-24. |
| Sherbrooke,
W.C. 2002. Phrynosoma cornutum. Nocturnal Nesting,
Eggs, Nest Predation, Hatchlings. Herpetol. Rev., 33(3):
206-208. |
| Sherbrooke,
W.C., E.R. Brown & J.L. Brown. 2002. Phrynosoma
hernandesi. Successful Open-Mouthed Threat Defense.
Herpetol. Rev., 33(3): 208. |
Sherbrooke, W.C. & S.K. Frost. 1989. Integumental
chromatophores of a color-change, thermoregulating lizard,
Phrynosoma modestum (Iguanidae; Reptilia). Amer.
Mus. Novitates, 2943: 1-14. |
| Sherbrooke,
W.C. & M.D. Greenfield. 2002. Phrynosoma hernandesi.
Defensive Hiss. Herpetol. Rev., 33(3): 208-209. |
| Sherbrooke,
W.C., B.E. Martin & C.H. Lowe. 1998. Geographic Distribution.
Phrynosoma ditmarsi. Herpetol. Rev., 29(2): 110-111. |
Sherbrooke, W.C. & R.R. Montanucci. 1988. Stone mimicry
in the round-tailed horned lizard, Phrynosoma modestum
(Sauria: Iguanidae). J. Arid Env., 14: 275-284. |
Sherbrooke, W.C. & R.B. Nagle. 1996. A dorsal intraepidermal
mechanoreceptor in horned lizards (Phrynosoma;
Phrynosomatidae; Reptilia). J. Morph., 228(2): 145-154. |
Smith, C.F. 1941. Birth of horned toads. Copeia, 1941(2):
114. |
Smith, H.M. 1934. Notes on some lizards of the genus Phrynosoma
from Mexico. Trans. Kansas Acad. Sci., 37: 286-297. |
Smith, H.M. 1942. Mexican herpetology miscellany. 2. A
new horned lizard from Durango. Proc. U.S. Natn. Mus.,
92: 349-395. |
| Smith,
H.M., K. Adler, D. Chiszar & F. van Breukelen. 1999.
Phrynosoma hernandesi: Correct Spelling. Herpetol.
Rev., 30(2): 74-76. |
Smith, H.M. & O.A. Flores Villela. 1994. Noteworthy
lizards of the genera Phrynosoma and Sceloporus
in the Museo de Zoologica "Alfonos L. Herrera".
Bull. Maryland Herpetol. Soc., 30(3): 114-119. |
Smith, H.M. & W. Reeve. 1951. The generic name of
the horned lizards. Herpetologica, 7(2): 53-55. |
Stejneger, L. 1906. A new lizard of the genus Phrynosoma
from Mexico. Proc. U.S. Natl. Mus., 29(1437): 565-567. |
Strecker, J.K. 1908. Notes on the breeding habits of Phrynosoma
cornutum and other Texas lizards. Proc. Biol. Soc.
Washington, 21: 165-170. |
Tanner, W.W. & J.E. Krogh. 1973. Ecology of Phrynosoma
platyrhinos at the Nevada test site, Nye County,
Nevada. Herpetologica, 29(4): 327-342. |
| Tollestrup,
K. 1981. The Social Behavior and Displays of Two Species
of Horned Lizards, Phrynosoma platyrhinos and
Phrynosoma coronatum. Herpetologica, 37(3): 130-141. |
| Turner,
D.S. & J. Rorabaugh. 1998. Phrynosoma platyrhinos.
Predation. Herpetol. Rev., 29(2): 101. |
Turner, F.B. & P.A. Medica. 1982. The distribution
and abundance of the flat-tailed horned lizard (Phrynosoma
mcalli). Copeia, 1982(4): 815-823. |
Tyler, J.D. 1977. Coachwhip preys on horned lizard. Southwestn.
Nat., 22(1): 146. |
Van Denburgh, J. 1894. Phrynosoma solaris, with
a note on its distribution. Proc. California Acad. Sci.,
2(4): 456. |
Van Devender, T.R. & C.W. Howard. 1973. Notes on natural
nests and hatching success in the regal horned lizard
(Phrynosoma solare) in southern Arizona. Herpetologica,
29(3): 238-239. |
| Wederkinch,
E. 1973. Tudseleguaner. NHF, 16(7): 130-136. |
Weese, A.O. 1917. An experimental study of the reactions
of the horned lizard, Phrynosoma modestum Girard,
a reptile of the semi-desert. Biol. Bull., 32(2): 98-116. |
Whitford, W.B. & W.G. Whitford. 1973. Combat in the
horned lizard, Phrynosoma cornutum. Herpetologica,
29(2): 191-192. |
Whitford, W.G. & M. Bryant. 1979. Behavior of a predator
and its prey: the horned lizard Phrynosoma cornutum
and harvester ants (Pogonomyrmex spp.). Ecology,
60(4): 686-694. |
| Whiting,
M.J. & J.R. Dixon. 1996. Phrynosoma modestum.
Cat. Amer. Amphib. Rept., 630: 1-6. |
Whiting, M.J., J.R. Dixon & R.C. Murray. 1993. Spatial
distribution of a population of Texas horned lizards (Phrynosoma
cornutum) (Phrynosomatidae) relative to habitat and
prey. Southwestn. Nat., 38(2): 150-154. |
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